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Quercus taxonomy
Quercus macrocarpa Michx.
EOL Text
Trees , deciduous, to 30(-50) m. Bark dark gray, scaly or flat-ridged. Twigs grayish or reddish, 2-4 mm diam., often forming extensive flat, radiating, corky wings, finely pubescent. Buds 2-5(-6) mm, glabrous. Leaves: petiole (6-)15-25(-30) mm. Leaf blade obovate to narrowly elliptic or narrowly obovate, often fiddle-shaped, (50-)70-150(-310) × (40-)50-130(-160) mm, base rounded to cuneate, margins moderately to deeply lobed, toothed, deepest sinuses near midleaf (at least in proximal 2/3), sinuses reaching nearly to midrib, longer lobes grading into shallow lobes or merely simple teeth distally, shallower, compound lobes proximally, secondary veins arched, divergent, 4-5(-10) on each side, apex broadly rounded or ovate; surfaces abaxially light green or whitish, with minute appressed-stellate hairs forming dense, rarely sparse, tomentum, erect felty hairs absent, adaxially dark green or dull gray, sparsely puberulent to glabrate. Acorns 1-3 on stout peduncle (0-)6-20(-25) mm; cup hemispheric or turbinate, (8-)15-50 mm deep × (10-)20-60 mm wide, enclosing 1/2-7/8 nut or more, scales closely appressed, laterally connate, broadly triangular, keeled, tuberculate, finely grayish tomentose, those near margins often with soft awns to 5-10 mm or more, forming fringe around nut; nut light brown or grayish, ovoid-ellipsoid or oblong, (15-)25-50 × (10-)20-40 mm, finely puberulent or floccose. Cotyledons distinct. 2 n = 24.
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Rights holder/Author | eFloras.org Copyright © Missouri Botanical Garden |
Source | http://www.efloras.org/florataxon.aspx?flora_id=1&taxon_id=233501058 |
Blue oak, mossycup oak, mossy-overcup oak
Bur oak on uplands is often associated with calcareous soils. In the "driftless" area of southwestern Wisconsin, it is commonly found on limestone ridges; in Kentucky, it is more prevalent on limestone soils than on soils derived from shales and sandstone (5). In western Iowa, it can be found as a dominant on soils of either limestone or sandstone origin. Throughout much of the prairie region of the Midwest, bur oak is found on droughty sandy plains, black prairie loams, and on loamy slopes of south and west exposure. Toward the western edge of its range, such as in eastern Kansas, it is more abundant on the more moist north-facing slopes than on southfacing slopes (2). Bur oak often dominates severe sites with thin soils, heavy claypan soils, gravelly ridges, and coarse-textured loessial hills. The predominant soil orders on which bur oak is found include Alfisols in the central and southern parts of its range, and Mollisols and Spodosols in the western and northern parts of its range, respectively.
Bur oak is also an important bottom-land species throughout much of its range. In the Central States Region and southward, it is found on moist flats and on hummocky bottoms. Northward, in southern Michigan, it has been found in high densities on slightly elevated ridges within wet bottom-land forests occupying old glacial lake beds and drainage ways (20).
Bur oak frequently forms a fringe between the prairie and upland forest in northern Illinois and eastern Iowa, notably at the outer edges of "breaks" and bluffs along streams and around limestone outcrops. It is a valuable timber species on favorable bottom-land sites within this region.
Within the Great Plains Region, it is frequently found in stream bottoms and stream terraces. In North Dakota, bur oak is a major component of the flood-plain forests of the Missouri River (11). Here it may predominate in old stands on high terraces near the edge of the flood plain. It is absent in low terraces near the center of the flood plain. Along adjacent draws and upper slopes, it becomes the first tree established along prairie edges. Bluffs along the Missouri River and its tributaries in eastern Nebraska are frequently covered with bur oaks that range in size from small trees near the base of bluffs to shrublike growth near the top.
In the Black Hills of western South Dakota and the Bear Lodge Mountains of northeastern Wyoming, bur oak grows at low elevations between the ponderosa pine forest and the grasslands (21). Here, it ranges in size from a shrub under a pine canopy at higher elevations to a tree up to 21 m (69 ft) tall along stream bottoms at lower elevations.
More info for the term: fresh
Conditions considered best for bur oak germination were not well documented in the available literature. In general, germination appears to decrease with desiccation and likely acorn age, but in its northern range, bur oak acorn germination requires 60 days or more of cold stratification ([248], review by [28]).
In controlled conditions, germination of bur oak acorns can be as high as 80%. After 25 to 45 days of alternating temperatures of 86 °F and 68 °F (30/20 °C) in the laboratory, bur oak germination averaged 45% (review [28]). Studies at a greenhouse container nursery showed that bur oak acorns germinated slowly and incompletely. Germination decreased as acorns dried. When acorns were 100%, 80%, and 65% of their fresh weight, germination was about 80%, 35%, and 0%, respectively. Bur oak acorns collected from North Dakota required 90 to 120 days of stratification before germinating. High temperatures were required for continued shoot growth [248].
A Forest Service nursery handbook reports that bur oak acorns have no dormancy and recommends planting acorns immediately follow harvest [268]. In Nebraska, 30% of bur oak seeds germinated within 1 month of falling (review by [125]).
Bur oak is monoecious; male and female flowers in separate catkins are home on the current year's branchlets. It flowers shortly after the leaves appear, from about the first of April in the southern part of its range to about mid-June in the north (5). Pollen from one tree appears to germinate better on the stigmas of another, favoring cross pollination.
Acorns of bur oak make up much of the food of red squirrels and are also eaten by wood ducks, white-tailed deer, New England cottontails, mice, thirteen-lined ground squirrels, and other rodents (5).
On coal-mine spoils with a pH of 5.6 in eastern Kansas, planted bur oak was one of the better performers of several tree species tested (7). After 22 years, it attained a mean height of 8.5 m (28 ft) and a d.b.h. of 12.2 cm (4.8 in). The species is also widely planted in shelterbelts because of its drought tolerance.
Bur Oak is a common tree that is found in every county of Illinois. Habitats include moist bottomland woodlands, upland woodlands, and savannas where deciduous trees are dominant. This tree is most commonly found in bottomland woodlands a little outside of the flood zone. It also occurs in savannas and can be an invader of prairies because of its resistance to fire. Occasionally, Bur Oak is cultivated as a landscape tree, where it can become quite large.
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Rights holder/Author | Copyright © 2002-2014 by Dr. John Hilty |
Source | http://www.illinoiswildflowers.info/trees/plants/bur_oak.htm |
Bur oak is one of the most drought resistant of the North American oaks. In the northwestern part of its range, the average annual precipitation is as low as 380 mm (15 in). Here, the average minimum temperature is 4° C (40° F), and the average growing season lasts only 100 days. To the south bur oak grows in areas having an average precipitation exceeding 1270 mm (50 in) per year, minimum temperatures of -7° C (20° F), and a growing season of 260 days. The best development of the bur oak occurs in southern Illinois and Indiana, where the average annual precipitation is about 1140 mm (45 in), minimum temperature is -29° C (-20° F), and the growing season is 190 days (5).
More info for the term: tree
Bur oak acorns are animal dispersed. Small mammals are the most likely dispersers. Birds are less likely dispersers, given the large size of bur oak acorns. Likelihood of bird dispersal may increase if only bur oak acorns are available or if small acorns are produced in a given season or on a given site. One study reported that bur oak acorns were dispersed by blue jays in Iowa (Johnson unpublished data [124]), but in a later Iowa study, blue jays avoided bur oak acorns [127].
Animal-mediated dispersal distances as great as 490 feet (150 m) were reported in forest fragments in southern Ontario. Rodents were the suspected dispersal agent. In this study, researchers searched a maximum distance of 490 feet (150 m) between seedlings and the nearest fruiting tree, suggesting that acorn dispersal distances may have been even greater than reported [110]. Near Manhattan, Kansas, fox squirrels cached bur oak acorns an average of 59 feet (18 m) from the source pile [228], but when researchers evaluated the number of seedlings occurring beyond the woodland edge in the same area, bur oak seedlings were found a maximum of 169 feet (51.5 m) from the forest edge [229].
Bur oak is a slow-growing tree (5). In 12- to 16-year-old plantations on Iowa upland sites, average annual height growth ranged from 0.09 to 0.52 m (0.3 to 1.7 ft) and diameter growth from less than 2.5 to 6.4 mm (0.1 to 0.25 in). In the shelterbelts of the northern Great Plains, an annual height growth of about 0.3 m (1 ft) was reported for trees kept under clean cultivation.
In Iowa, 10-year d.b.h. growth of bur oak averaged 3.0 cm (1.2 in) for 10- to 20-cm (4- to 8-in) trees, 3.6 cm (1.4 in) for 25- to 36-cm (10- to 14-in) trees, 4.6 cm (1.8 in) for 41- to 51-cm (16- to 20-in) trees, and 5.6 cm (2.2 in) for trees 56 cm (22 in) and larger. More rapid growth has been reported in Kansas where trees 35 to 40 years old averaged 2.5 cm (I in) growth in d.b.h. in 3.8 years. Approximately the same growth rate has been observed in the northern Mississippi Delta region.
Bur oak is said to have reached a height of 52 m (170 ft) and a d.b.h. of 213 cm (84 in) in the lower Ohio Valley. On the better sites, mature trees generally grow 24 to 30 in (80 to 100 ft) tall, 91 to 122 cm (36 to 48 in) in d.b.h., and live 200 to 300 years. Characteristically, they have a massive, clear trunk and a broad, open crown of stout branches.
In the oak openings of southern Wisconsin and in the prairie border areas to the south and west, bur oak often is found in nearly pure stands (3,5). The trees are widely spaced, short-boled, and often uniform in size. Trees in a 50- to 65-year-old stand in eastern Nebraska were 9 to 12 in (30 to 40 ft) tall and spaced at intervals of 3 to 12 in (10 to 40 ft). Bur oak grows 21 in (70 ft) tall on the fertile soils in this region, but on dry, limestone ridges, the trees may be less than 7.6 in (25 ft) tall at 150 years of age. In Minnesota, bur oak is short lived on the poorer sites.
Timber volumes in the bur oak type of Iowa were estimated to be 15.4 m³/ha (1,100 fbm/acre), three-fourths of which were bur oak.